Express Newspapers was the only publisher to post a rise in its daily title circulation. The Daily Star was up 20% year on year to 887,106. The circulation of its Sunday title rose 6.6%.
Guardian Media Group's guardian.co.uk site increased its traffic by 43% year on year, with a daily average of 1.5m, but failed to overtake the MailOnline. The Guardian's July print circulation was down 2.7% year on year to 328,773. The Observer was down by 7%.
Independent News & Media suffered the biggest decrease in circulation for July across its national titles. The Independent and Independent on Sunday were both down by 20%. Traffic to the newspaper's website was up 29% year on year to a daily average of 386,154 users.
The Sun remained the biggest daily in Britain with a circulation of 3.1m. Fellow News International title The Times was down 5% year on year to 580,483, while The Sunday Times was up 3% to 1.2m.
The Mirror Group recorded the biggest spike in online traffic, increasing its average daily users by 81% year on year to 495,798.